This module examines how biology can be explored at scales from individual signalling pathways to whole organisms using methods and approaches from modern molecular systems biology. ‘Omics approaches including proteomics, genomics, transcriptomics and metabolomics are taught through the lens of these different experimental questions. Throughout, students will evaluate the field of molecular systems biology, investigate landmark papers using these different methods across different scales, and appreciate and the extent to which different investigative approaches and concepts are complementary to each other.